New strikes are causing more disruption for rail users in the last 24 hours of release on the security.
RMT union members in the North, in the South-West of the Railway, Merseyrail and Greater Anglia to take measures for the third time in less than a week.
The union said the demolition of the guards and the introduction of driver only operates trains of risks to the safety of the passengers.
But the railroad companies have insisted on the fact that the movement is safe and is already widely used.
Picket lines appeared at a few railway stations affected by the strike, and passengers are faced with delays, cancellations and replacement buses in some parts of the country.
All four operators said some services would still be running:
North said that more than 1,300 trains would run between 07:00 and 19:00 GMT, representing approximately 50% of his calendar of the week
Greater Anglia said it planned to keep conductors on the trains, and would be operating a normal service
Merseyrail said no trains would run on the Kirkby, Ellesmere Port, or Hunts Cross lines, but most of the others would work between 07:00 and 19:00
South-West of the Railway, which operates particularly in the areas of London Waterloo, Reading, Exeter and Southampton, said he planned to run for more than 70% of its calendar week and of longer trains to provide more seats. The Island Line will work normally
RMT general secretary Mick Cash said: “It is frankly ridiculous that we have been able to negotiate long-term agreements in Scotland and Wales which protect the guards and the safety of the passengers, but we were denied the same opportunities with the railway companies in England.
“This suspension of the normal industrial relations by employers has to end if we are to make progress towards a solution that ensures the safety of train travel for all.
“RMT is ready for talks in each of these disputes.”
A Department of Transport spokesman said: “This is a dispute between a private company and the RMT.
“However, the secretary of state for transport recognises the disruption caused by passengers and met with the leaders of the union, on several occasions, including recently, in December, to help put an end to the strike.
“Nobody is losing their job because of the driver controlled the operation of trains, employees have been guaranteed jobs and wages for several years.”